A COLEOPTERIST’S COLOUR-BY-NUMBER
Hand-tinted plate of Otiorhynchus ovatus. For hand-tinting, color each area in the drawing by the number
in that area, using the following color code:
1: brown
David R. Maddison is professor and curator in the Department of Entomology at the University of Arizona. His research focuses on the evolution of beetles, and developing methods and software for phylogenetic analysis.
